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Moqueguahippus Temporal range: Late Oligocene (Deseadan) ~28.4–23.03 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Notoungulata
Family:	†Notohippidae
Genus:	†Moqueguahippus Shockey et al. 2006
Species:	†M. glycisma
Binomial name
†Moqueguahippus glycisma Shockey et al., 2006

Moqueguahippus is an extinct genus of notohippid notoungulates that lived during the Late Oligocene of what is now Peru. Fossils of this genus have been found in the Moquegua Formation of Peru, which it was named after.

Etymology

The genus name, Moqueguahippus, refers to the Moquegua Formation where it was discovered, and -hippus meaning "horse", an epithet used to describe most notohippids. The specific name, glycisma, is derived from the Greek word Glykisma, meaning "cake", in reference to the type locality, Pan de Azucar, meaning "Sugarloaf Mountain".

Description

Moqueguahippus is a relatively large notohippid. Its skull was rather massive and had a short muzzle. The dentition was complete and continuous, lacking a diastema, and the premolars and molars were high-crowned (hypsodont), like in its relative Rhynchippus. The premolars had a well developed cingulum, while the molars were elongated. The shape of the premolars and molars was reminiscent to those of Equidae. Moqueguahippus, like its relatives, may have had a robust body with slender limbs. Its most peculiar distinctive characteristics were the cup-shaped internal cingula and the lack of cingula on the premolars, and a thick layer of cementum.

Taxonomy

Moqueguahippus was first described in 2006, based on fossilized remains found in the Moquegua Formation in Peru, in deposits dating to the Late Oligocene. Moqueguahippus was placed within the family Notohippidae, a family of toxodont notoungulates with horse-like teeth, with very high-crowned molars. More recent research, however, suggests that this group is paraphyletic, containing increasingly derived forms, some of them close to the ancestors of "true" Toxodontidae. In 2021, it was found to be a derived notohippid, being the sister taxon of a clade including the Miocene genera Argyrohippus and Notohippus.

The following position of the Toxodontia is based on Martínez et al. 2021, showing the position of Moqueguahippus.

References

1. ^ Shockey, Bruce J.; Salas, Rodolfo; Quispe, Rossana; Flores, Alexander; Sargis, Eric J.; Acosta, Jorge; Pino, Adan; Jarica, Nestor J.; Urbina, Mario (2006-03-30). "Discovery of Deseadan fossils in the upper Moquegua Formation (late Oligocene—?early Miocene) of southern Peru". Journal of Vertebrate Paleontology. 26 (1): 205–208. doi:10.1671/0272-4634(2006)26[205:DODFIT]2.0.CO;2. ISSN 0272-4634. S2CID 85593383.
2. Shockey, Bruce J.; Gismondi, Rodolfo Salas; Gans, Phillip; Jeong, Annie; Flynn, John J. (2009). "Paleontology and Geochronology of the Deseadan (late Oligocene) of Moquegua, Perú". American Museum Novitates (3668): 1–24. doi:10.1206/662.1. ISSN 0003-0082.
3. Cerdeño, Esperanza; Vera, Bárbara (2014-06-07). "New data on diversity of Notohippidae from the Oligocene of Mendoza, Argentina". Journal of Vertebrate Paleontology. 34 (4): 941–950. Bibcode:2014JVPal..34..941C. doi:10.1080/02724634.2014.837054. hdl:11336/32354. ISSN 0272-4634. S2CID 85991987.
4. ^ Martínez, Gastón; Dozo, María T.; Gelfo, Javier N.; Ciancio, Martín R.; González-José, Rolando (2021-02-11). "A new toxodont (Mammalia, Panperissodactyla, Notoungulata) from the Oligocene of Patagonia, Argentina, and systematic considerations on the paraphyletic 'Notohippidae'". Journal of Systematic Palaeontology. 18 (24): 1995–2013. doi:10.1080/14772019.2021.1872723. ISSN 1477-2019. S2CID 232116246.

v t e Genera of South American native ungulates
Kingdom: Animalia Phylum: Chordata Class: Mammalia Superorder: Laurasiatheria
Xenungulata, Pyrotheria, Astrapotheria, and other minor groups †Kollpaniidae? Andinodus Molinodus Pascualodus? Pucanodus Simoclaenus Tiuclaenus †Didolodontidae Didolodus Enneoconus Ernestokokenia Escribania? Lamegoia Paulogervaisia Raulvaccia Ricardocifellia Saltaodus Umayodus Xesmodon †Protolipternidae Asmithwoodwardia Miguelsoria Protolipterna †Xenungulata †Carodniidae Carodnia Etayoa Notoetayoa Rodcania †Pyrotheria †Colombitheriidae Colombitherium Proticia †Pyrotheriidae Baguatherium Berracotherium Carolozittelia Griphodon Propyrotherium Pyrotherium †Astrapotheria Eoastrapostylops †Trigonostylopidae Shecenia Tetragonostylops Trigonostylops †Astrapotheriidae Antarctodon Astrapodon Comahuetherium Liarthrus Maddenia Parastrapotherium †Albertogaudryinae Albertogaudrya Astraponotus †Astrapotheriinae Astrapothericulus Astrapotherium Scaglia †Uruguaytheriinae Granastrapotherium Hilarcotherium Uruguaytherium Xenastrapotherium
Litopterna †Eulitopterna †Adianthidae Adianthus Proadiantus Proectocion? Proheptaconus Thadanius Tricoelodus †Macraucheniidae Polymorphis †Cramaucheniinae Caliphrium Coniopternium Cramauchenia Llullataruca Phoenixauchenia Pternoconius Theosodon †Macraucheniinae Cullinia Huayqueriana Macrauchenia Macraucheniopsis Micrauchenia Oxyodontherium Paranauchenia Promacrauchenia Scalabrinitherium Windhausenia Xenorhinotherium †Proterotherioidea †Anisolambdidae †Anisolambdinae Anisolambda Eolicaphrium Guilielmofloweria Lambdaconops Paranisolambda Polyacrodon Protheosodon Wainka †Sparnotheriodontinae Heteroglyphis Notiolofos Phoradiadius Sparnotheriodon Victorlemoinea †Proterotheriidae Mesolicaphrium Neodolodus Olisanophus Promylophis Prothoatherium Pseudobrachytherium Salladolodus †Megadolodinae Bounodus Megadolodus †Proterotheriinae Anisolophus Brachytherium Diadiaphorus Diplasiotherium Eoauchenia Epecuenia Epitherium Lambdaconus Neobrachytherium Neolicaphrium Paramacrauchenia Picturotherium Prolicaphrium Proterotherium Tetramerorhinus Thoatheriopsis Thoatherium Uruguayodon Villarroelia †Notopterna †Amilnedwardsiidae Amilnedwardsia Ernestohaeckelia Rutimeyeria †Indaleciidae Adiantoides Indalecia †Notonychopidae Notonychops Requisia Incertae sedis Kyraodus Proacrodon
Notoungulata incertae sedis Acoelodus Archaeogaia Carolodarwinia Perutherium †Notioprogonia Satshatemnus Seudenius †Henricosborniidae Henricosbornia Nanolophodon Othnielmarshia Peripantostylops Simpsonotus †Notostylopidae Boreastylops Chilestylops Edvardotrouessartia Homalostylops Notostylops Otrhonia Parastylops †Toxodontia †Homalodotheriidae Asmodeus Chasicotherium Homalodotherium Trigonolophodon †Isotemnidae Anisotemnus Coelostylodon Distylophorus Hedralophus Isotemnus Lophocoelus Pampatemnus Periphragnis Pleurocoelodon Pleurostylodon Rhyphodon Thomashuxleya †Eutoxodontia †Leontiniidae Anayatherium Ancylocoelus Colpodon Coquenia Elmerriggsia Gualta Henricofilholia Huilatherium Leontinia Martinmiguelia Purperia? Scarrittia Taubatherium Termastherium †Notohippidae Acoelohyrax Argyrohippus Deuterotherium? Edvardocopeia Eomorphippus Eurygenium Mendozahippus Moqueguahippus Morphippus Nesohippus Notohippus Pampahippus Pascualihippus Patagonhippus Perhippidion Plexotemnus Puelia Purperia? Rhynchippus Rosendo Stilhippus Teushentherium Trimerostephanos †Toxodontidae †Nesodontinae Adinotherium Nesodon Proadinotherium †Toxodontinae Abothrodon Andinotoxodon Calchaquitherium Ceratoxodon Charruatoxodon Dinotoxodon Eutomodus Falcontoxodon Gyrinodon Haplodontherium Hemixotodon Hoffstetterius Hyperoxotodon Mesenodon Mesotoxodon Minitoxodon Mixotoxodon Neoadinotherium Neotoxodon Nesodonopsis Nonotherium Ocnerotherium Palyeidodon Paratrigodon Pericotoxodon Piauhytherium Pisanodon Plesiotoxodon Posnanskytherium Prototrigodon Stenotephanos Stereotoxodon Toxodon Toxodontherium Trigodon Trigodonops Xotodon †Typotheria Campanorco Griphotherion †Archaeopithecidae Archaeopithecus Teratopithecus †Oldfieldthomasiidae Allalmeia Brachystephanus Camargomendesia Colbertia Dolichostylodon Itaboraitherium Kibenikhoria Maxschlosseria Oldfieldthomasia Paginula Suniodon Tsamnichoria Ultrapithecus Xenostephanus †Interatheriidae Antofagastia Ignigena Johnbell Munizia Punapithecus †Interatheriinae Archaeophylus Argyrohyrax Boleatherium Brucemacfaddenia Caenophilus Choichephilum Cochilius Eopachyrucos Federicoanaya Interatherium Juchuysillu Miocochilius Neoicochilus Patriarchus Proargyrohyrax Progaleopithecus Protypotherium Santiagorothia †Notopithecinae Antepithecus Guilielmoscottia Notopithecus Transpithecus †Mesotheriidae Fiandraia? Trachytherus †Mesotheriinae Altitypotherium Caraguatypotherium Eotypotherium Eutypotherium Hypsitherium Mesotherium Microtypotherium Plesiotypotherium Pseudotypotherium Rusconitherium Typotheriopsis †Archaeohyracidae Archaeohyrax Archaeotypotherium Bryanpattersonia Eohyrax Pascualhyrax Protarchaeohyrax Pseudhyrax Punahyrax †Hegetotheriidae †Hegetotheriinae Ethegotherium Hegetotheriopsis Hegetotherium Hemihegetotherium Prohegetotherium Pseudohegetotherium Sallatherium †Pachyrukhinae Medistylus Propachyrucos Prosotherium Tremacyllus †Pachyrukhini Pachyrukhos Paedotherium Tremacyllus

• Toxodonts
• Prehistoric placental genera
• Oligocene mammals of South America
• Paleogene Peru
• Fossils of Peru
• Deseadan
• Fossil taxa described in 2006