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Stereognathus Temporal range: Bathonian-Barremian?, 167.7–165.7 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
The postcanine tooth of Stereognathus ooliticus, from Owen 1857.
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Clade:	Therapsida
Clade:	Cynodontia
Family:	†Tritylodontidae
Genus:	†Stereognathus Charlesworth, 1854
Type species
Stereognathus ooliticus Charlesworth, 1854
Potential species
S. chuannanensis (He & Cai, 1984; originally Polistodon) S. kolossovi (Lopatin & Agadjanian, 2008; originally Xenocretosuchus) S. kuwajimaensis (Matsuoka et al., 2016; originally Montirictus) S. sibiricus (Tatarinov & Mashenko, 1999; originally Xenocretosuchus)
Synonyms
S. hebridicus? Waldman & Savage, 1972

Stereognathus is an extinct genus of tritylodontid cynodonts from the Middle Jurassic of the United Kingdom. There is a single named species: S. ooliticus, named after the Great Oolite deposits of England. A second species, S. hebridicus, was named after the Hebrides in Scotland, where it was found; it was synonymized with S. ooliticus in 2017.

Discovery and naming

S. ooliticus was the first tritylodontid species ever found, being described by Charlesworth in 1855 and later by Sir Richard Owen. In 1972, S. hebridicus was named from several postcanine teeth recovered in Bathonian age deposits on the Isle of Skye in Scotland by palaeontologists R. J. G. Savage and Michael Waldman. Despite being the first tritylodontid genus found and named, Stereognathus remains poorly represented, being known mainly from isolated molar teeth. There is, however, one holotype fragment of maxilla with three damaged molars, and a second fragment of maxilla with four sets of molar roots.

In 2017, Elsa Panciroli and colleagues found no points of variance between the anatomy of S. ooliticus and S. hebridicus, accounting for changes with growth, which makes S. hebridicus a junior synonym of S. ooliticus. Also in 2017, Alexander Averianov and colleagues considered species of Polistodon, Xenocretosuchus, and Montirictus to be within the genus Stereognathus; they also retained S. hebridicus as a separate species. However, this would mean Stereognathus persisted almost unchanged for over 60 million years across the whole of Eurasia, which would be very unusual. The issue remains unresolved.

References

1. Fossilworks. "Stereognathus." Retrieved from
2. ^ Panciroli, E.; Walsh, S.; Fraser, N.C.; Brusatte, S.L.; Corfe, I. (2017). "A reassessment of the postcanine dentition and systematics of the tritylodontid Stereognathus (Cynodontia, Tritylodontidae, Mammaliamorpha), from the Middle Jurassic of the United Kingdom". Journal of Vertebrate Paleontology. 37 (5): e1351448. doi:10.1080/02724634.2017.1351448. hdl:20.500.11820/b140e841-1ad5-4cbe-aad7-7bb588f9f0c1.
3. Charlesworth, E 1855. Reprint British Association. 1854 Liverpool Abstracts, 80.
4. Owen, R 1856 On the affinity of Stereognathus ooliticus (Charlesworth) a mammal from the Oolitic slate of Stonesfield . Quarterly Journal of the Geological Society of London, 13, 1-11.
5. Waldman, M and Savage, R.J.G 1972 The first Jurassic mammal from Scotland. Journal of the Geological Society of London 128:119-125
6. Averianov, A.O.; Martin, T.; Lopatin, A.V.; Schultz, J.A.; Skutschas, P.P.; Rico, S.; Krasnolutskii, S.A. (2017). "A tritylodontid synapsid from the Middle Jurassic of Siberia and the taxonomy of derived tritylodontids". Journal of Vertebrate Paleontology. 37 (5): e1363767. doi:10.1080/02724634.2017.1363767.

v t e Cynodontia
Kingdom: Animalia Phylum: Chordata Clade: Synapsida Clade: Therapsida Clade: Eutheriodontia Synapsida see Synapsida Cynodontia see below↓
Cynodontia Cynodontia †Dvinia †Nanocynodon †Madysaurus †Uralocynodon †Charassognathidae Abdalodon Charassognathus Nshimbodon †Procynosuchidae Procynosuchus Sludica Epicynodontia †Bolotridon †Cynosaurus †Nanictosaurus †Nythosaurus †Platycraniellus †Thrinaxodon †Vetusodon †Galesauridae Galesaurus Progalesaurus Eucynodontia †Kataigidodon †Cynognathia see Cynognathia Probainognathia see below↓
Probainognathia Probainognathia †Agudotherium †Candelariodon †Charruodon †Lumkuia †Protheriodon †Tessellatia †Chiniquodontidae Aleodon Chiniquodon Cromptodon? Riojanodon †Probainognathidae Bonacynodon Probainognathus †Ecteniniidae Diegocanis Ecteninion Trucidocynodon Prozostrodontia †Alemoatherium †Deccanodon †Dromatherium †Inditherium †Microconodon †Polonodon †Pseudotriconodon †Rewaconodon †Santacruzgnathus †Therioherpeton †Tricuspes †Prozostrodontidae Prozostrodon Pseudotherium †Tritheledontidae Chaliminia Diarthrognathus Elliotherium Irajatherium Pachygenelus Riograndia? Tritheledon Mammaliamorpha †Adelobasileus †Botucaraitherium †Brasilodon †Sinoconodon †Tikiodon †Tritylodontidae Bienotherium Bienotheroides Bocatherium Dianzhongia Dinnebitodon Fossiomanus Kayentatherium Lufengia Montirictus Nuurtherium Oligokyphus Polistodon Shartegodon Stereognathus Tritylodon Xenocretosuchus Yuanotherium Yunnanodon Mammaliaformes see below↓
Mammaliaformes Mammaliaformes †Bocaconodon †Brachyzostrodon †Bridetherium †Cherwellia †Cifellilestes †Delsatia †Dianoconodon †Dinnetherium †Dyskritodon †Eozostrodon †Erythrotherium †Gondwanadon †Hadrocodium †Hallautherium †Helvetiodon †Indotherium †Indozostrodon †Megazostrodon †Morganucodon †Paceyodon †Paikasigudodon †Purbeckodon †Rosierodon †Sinoconodon? †Storchodon †Stylidens †Wareolestes †Woutersia †Kuehneotheriidae Fluctuodon Kotatherium Kuehneon Kuehneotherium †Shuotheriidae Feredocodon Itatodon? Paritatodon? Pseudotribos Shuotherium †Docodonta Agilodocodon Borealestes Castorocauda Cyrtlatherium Dobunnodon Docodon Docofossor Dsungarodon Ergetiis Gondtherium? Haldanodon Hutegotherium Itatodon? Khorotherium Krusatodon Microdocodon Paritatodon? Peraiocynodon Sibirotherium Simpsonodon Tashkumyrodon Tegotherium †Haramiyida Avashishta? Hypsiprymnopsis? Haramiyavia Kalaallitkigun Kollikodon? Mojo? Theroteinus Thomasia Euharamiyida Allostaffia? Arboroharamiya Baidabatyr? Butlerodon Cryoharamiya Kermackodon? Maiopatagium Megaconus? Qishou Sharypovoia Shenshou Sineleutherus Vilevolodon Woodeatonia Xianshou Gondwanatheria? Hahnodontidae? Cifelliodon Denisodon Hahnodon Crown-Mammalia see Mammalia
See also Incertae sedis †Cistecynodon †Eoraetia †Gaumia †"Hahnia" †Kunminia †Lepagia †Meurthodon †Mitredon †Panchetocynodon †Redondagnathus †Saurodesmus? Paraphyletic / Polyphyletic groups Dromatheriidae Morganucodonta Symmetrodonta Triconodonta
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