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Coraciiformes Temporal range: Middle Eocene to present PreꞒ Ꞓ O S D C P T J K Pg N
European roller Coracias garrulus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Clade:	Picodynastornithes
Order:	Coraciiformes Forbes, 1884
Families
Meropidae Brachypteraciidae Coraciidae Todidae Momotidae Alcedinidae For prehistoric taxa, see text.
Global distribution of the kingfisher and allies.

The Coraciiformes /kɒrəˈsaɪ.ɪfɔːrmiːz/ are a group of usually colourful birds including the kingfishers, the bee-eaters, the rollers, the motmots, and the todies. They generally have syndactyly, with three forward-pointing toes (and toes 3 & 4 fused at their base), though in many kingfishers one of these is missing. The members of this order are linked by their "slamming" behaviour, thrashing their prey onto surfaces to disarm or incapacitate them.

In the past, Coraciiformes encompassed all anisodactyl (including syndactyl) members of the Cavitaves. This means that the birds currently classified in Leptosomiformes (cuckoo-roller) and Bucerotiformes (hornbills, hoopoes and allies) were formerly classified in Coraciiformes.

This is largely an Old World order, with the representation in the New World limited to the dozen or so species of todies and motmots, and a mere handful of the more than a hundred species of kingfishers.

The name Coraciiformes means "raven-like". Specifically, it comes from the Latin language "corax", meaning "raven" and Latin "forma", meaning "form".

Systematics

The phylogenetic relationships between the six families in the order Coraciiformes is shown below. The cladogram is based on a large study by Richard Prum and colleagues published in 2015. The number of species in each family is taken from the list maintained by Frank Gill, Pamela C. Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC).

Several extinct coraciiform families are only known from Paleogene fossils. They probably belong to the basal group and are sometimes difficult to assign because they were even closer still to the Piciformes (see also Neanis). In addition, there are some prehistoric genera which are likewise difficult to place into a family. At least the Eocoraciidae are very basal, but the Late Eocene (some 35 mya) Geranopteridae form a superfamily Coracioidea with the extant rollers and ground-rollers already (Mayr & Mourer-Chauviré 2000). A few prehistoric taxa of the present-day families have been described; see the family articles for details.

Taxonomic sequence

• 

  Unresolved
  • Genus Quasisyndactylus (fossil; Middle Eocene of Messel, Germany) - alcediniform, basal?
  • Genus Cryptornis (fossil; Late Eocene of France) – bucerotid? geranopterid?
  • Family Primobucconidae (fossil), including Primobucco and Septencoracias
  • Coraciiformes gen. et spp. indet. PQ 1216, QU 15640 (fossil; Late Eocene of Quercy, France: Mayr & Mourer-Chauviré 2000)
  • Genus Protornis (fossil: Oligocene of Switzerland) – basal to motmotids and meropids?

A recent study suggest that the following families may belong to a separate order called Bucerotiformes. The results still in dispute though.

• Family Bucorvidae (ground hornbills)
• Family Bucerotidae (hornbills)
• Family Upupidae (hoopoe)
• Family Phoeniculidae (woodhoopoes)

The Leptosomidae (cuckoo roller) probably do not belong here. The trogons are sometimes placed here as a family Trogonidae. The Late Eocene Palaeospizidae are sometimes also placed in the Coraciiformes, as are the Early to Middle Eocene Primobucconidae and the Middle Eocene to Early Oligocene Sylphornithidae. The Primobucconidae at least indeed seem to belong here.

• Basal group
  • Family Eocoraciidae (fossil; Middle Eocene of Messel, Germany)
  • Family Geranopteridae (fossil; Late Eocene of Quercy, France – Early Miocene of Czech Republic) - includes "Nupharanassa" bohemica
  • Family Coraciidae (rollers)
  • Family Brachypteraciidae (ground-rollers)
  • Family Meropidae (bee-eaters)

• Suborder Alcedines
  • Family Todidae (todies)
  • Family Momotidae (motmots)
  • Family Alcedinidae (kingfishers)

See also

• List of Coraciiformes by population

References

1. "Coraciiformes". Britannica. Retrieved 13 September 2022.
2. Clements, J. F.; Schulenberg, T. S.; Iliff, M. J.; Roberson, D.; Fredericks, T. A.; Sullivan, B. L.; Wood, C. L. (2014). "The eBird/Clements checklist of birds of the world: Version 6.9". Archived from the original on December 17, 2014.
3. Terres, John K. (1980). The Audubon Society encyclopedia of North American birds (1st ed.). New York: A. A. Knopf. p. 104. ISBN 0-394-46651-9.
4. Prum, R.O.; Berv, J.S.; Dornburg, A.; Field, D.J.; Townsend, J.P.; Lemmon, E.M.; Lemmon, A.R. (2015). "A comprehensive phylogeny of birds (Aves) using targeted next-generation DNA sequencing". Nature. 526 (7574): 569–573. Bibcode:2015Natur.526..569P. doi:10.1038/nature15697. PMID 26444237.
5. Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (December 2023). "IOC World Bird List Version 14.1". International Ornithologists' Union. Retrieved 17 June 2024.
6. "Bucerotiformes". tolweb.org. Retrieved 31 March 2018.

• Johansson, Ulf S. & Ericson, Per G. P. (2003): Molecular support for a sister group relationship between Pici and Galbulae (Piciformes sensu Wetmore 1960). J. Avian Biol. 34(2): 185–197. doi:10.1034/j.1600-048X.2003.03103.x PDF fulltext
• Mayr, Gerald & Mourer-Chauviré, Cécile (2000): Rollers (Aves: Coraciiformes. s.s.) from the Middle Eocene of Messel (Germany) and the Upper Eocene of the Quercy (France). J. Vertebr. Paleontol. 20(3): 533–546. DOI:10.1671/0272-4634(2000)0202.0.CO;2 PDF fulltext
• Terres, John K. (1980) The Audubon Society Encyclopedia of North American Birds. ISBN 0-394-46651-9

External links

• Order Coraciiformes - Biodiversity Overview: Untamed Science
• Tree of Life: Coraciiformes

v t e Genera of trogons, hornbills, kingfishers, woodpeckers and their extinct allies
Kingdom: Animalia Phylum: Chordata Class: Aves Infraclass: Neognathae Clade: Coraciimorphae Clade: Cavitaves
Cavitaves Leptosomiformes Leptosomidae Leptosomus †Plesiocathartes? †Waltonavis Eucavitaves Trogoniformes Trogonidae Apalharpactes Apaloderma †Eotrogon Euptilotis †Foshanornis Harpactes †Masillatrogon †Paratrogon Pharomachrus †Primotrogon Priotelus †Septentrogon Trogon Picocoraciae Bucerotiformes See below ↓ Picodynastornithes Coraciiformes See below ↓ Piciformes See below ↓
Bucerotiformes Bucerotiformes †Laurillardiidae Laurillardia †Messelirrisoridae Messelirrisor Upupi Phoeniculidae †Phirriculus Phoeniculus Rhinopomastus Upupidae Upupa Buceroti Bucorvidae Bucorvus †Euroceros Bucerotidae Aceros Anorrhinus Anthracoceros Berenicornis Buceros Bycanistes Ceratogymna Horizocerus Lophoceros Ocyceros Penelopides Rhabdotorrhinus Rhinoplax Rhyticeros Tockus
Coraciiformes incertae sedis †Cryptornis †Paracoracias †Quasisyndactylus †Eocoraciidae Eocoracias †Geranopteridae Geranopterus †Primobucconidae Primobucco Septencoracias Brachypteraciidae Atelornis Brachypteracias Geobiastes Uratelornis Coraciidae †Chehuenia Coracias Eurystomus †Miocoracias Meropidae Meropogon Merops Nyctyornis Alcedines Alcedinidae Alcedininae Alcedo Ceyx Corythornis Ispidina Cerylinae Ceryle Chloroceryle Megaceryle Halcyoninae Actenoides Caridonax Cittura Clytoceyx Dacelo Halcyon Lacedo Melidora Pelargopsis Syma Tanysiptera Todiramphus Momotidae Aspatha Baryphthengus Electron Eumomota Hylomanes Momotus †Protornis? Todidae †Palaeotodus Todus
Piciformes incertae sedis †"Homalopus" †Pristineanis †Gracilitarsidae Gracilitarsus Neanis †Sylphornithidae Oligosylphe Palaegithalus? Sylphornis Galbuli †Jacamatia Bucconidae Bucco Chelidoptera Hapaloptila Hypnelus Malacoptila Micromonacha Monasa Nonnula Notharchus Nystalus Galbulidae Brachygalba Galbalcyrhynchus Galbula Jacamaralcyon Jacamerops Pici incertae sedis †Rupelramphastoides †Miopiconidae Miopico †Picavidae Picavus Ramphastides Capitonidae Capito †Capitonides Eubucco Lybiidae Buccanodon Cryptolybia Gymnobucco Lybius Pogoniulus Pogonornis Stactolaema Trachyphonus Tricholaema Megalaimidae Caloramphus Psilopogon Ramphastidae Andigena Aulacorhynchus Pteroglossus Ramphastos Selenidera Semnornithidae Semnornis Picides Indicatoridae Indicator Melichneutes Melignomon Prodotiscus Picidae See below ↓
Picidae incertae sedis †Bitumenpicus †Breacopus †Palaeonerpes †Palaeopicus †Piculoides Jynginae Jynx Picumninae Picumnus Sasiinae Sasia Verreauxia Picinae †Australopicus †Palaeonerpes †Pliopicus Nesoctitini Nesoctites Hemicircini Hemicircus Picini Campethera Celeus Chrysophlegma Colaptes Dinopium Dryocopus Gecinulus Geocolaptes Micropternus Meiglyptes Mulleripicus Pardipicus Piculus Picus Campephilini Blythipicus Campephilus Chrysocolaptes Reinwardtipicus Melanerpini Chloropicus Dendrocopos Dendrocoptes Dendropicos Dryobates Leiopicus Leuconotopicus Melanerpes Picoides Sphyrapicus Veniliornis Xiphidiopicus Yungipicus

• Coraciiformes
• Extant Eocene first appearances
• Bird orders
• Eocene taxonomic orders
• Oligocene taxonomic orders
• Miocene taxonomic orders
• Pliocene taxonomic orders
• Pleistocene taxonomic orders
• Holocene taxonomic orders